Walther Flemming

German anatomist Walther Flemming  recognized and explored the fibrous network within the nucleus, which he termed chromatin, or “stainable material.” (Flemming had actually discovered the chromosome, although the term would be coined a few years later by Heinrich Waldeyer.) Flemming noted that during cell division, the chromatin formed threadlike bodies, which he termed mitosen, from the Greek word for thread. Based on many observations of cells in various stages of division (Figure below), Flemming correctly deduced the sequence of chromosome movements during mitosis, which would be confirmed decades later by microscopy of live dividing cells. (from nature.com)

Drawing of Mitosis by Walther Flemming (link) from nature.com

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